1. rTG v1.0.4 : methods to analyse seasonal radial Tree Growth dataJernej Jevšenak, software Abstract: Methods for comparing different regression algorithms for describing the temporal dynamics of secondary tree growth (xylem and phloem). Users can compare the accuracy of the most common fitting methods usually used to analyse xylem and phloem data, i.e., Gompertz function, Double Gompertz function, General Additive Models (GAMs); and an algorithm newly introduced to the field, i.e., Bayesian Regularised Neural Networks (brnn). The core function of the package is XPSgrowth(), while the results can be interpreted using implemented generic S3 methods, such as plot() and summary(). Published in DiRROS: 24.11.2025; Views: 319; Downloads: 138
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2. MLFS v0.4.3 : Machine Learning Forest SimulatorJernej Jevšenak, software Abstract: Climate-sensitive, single-tree forest simulator based on data-driven machine learning. It simulates the main forest processes— radial growth, height growth, mortality, crown recession, regeneration, and harvesting—so users can assess stand development under climate and management scenarios. Published in DiRROS: 24.11.2025; Views: 284; Downloads: 162
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3. DendroTools v1.2.15 : linear and nonlinear methods for analyzing dendroclimatological dataJernej Jevšenak, software Abstract: Provides novel dendroclimatological methods, primarily used by the Tree-ring research community. There are four core functions. The first one is daily_response(), which finds the optimal sequence of days that are related to one or more tree-ring proxy records. Similar function is daily_response_seascorr(), which implements partial correlations in the analysis of daily response functions. For the enthusiast of monthly data, there is monthly_response() function. The last core function is compare_methods(), which effectively compares several linear and nonlinear regression algorithms on the task of climate reconstruction. Published in DiRROS: 24.11.2025; Views: 319; Downloads: 133
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4. Data from 'Spatiotemporal variability of dendroecological indicators in pedunculate oak (Quercus robur L.) tree-rings across Europe in relation to species distribution models' : version v1.0Andrei Popa, Jernej Jevšenak, Marcin K. Dyderski, Radosław Puchałka, Allan Buras, Ionel Popa, Martin Wilmking, Aleksandra Kalisty, Marcin Jakubowski, Eric Andreas Thurm, 2025, complete scientific database of research data Abstract: Climate is a primary, but non-stationary, driver of tree growth. Climate change is altering the sensitivity of forest growth to water availability and temperature over time. It is considered that pedunculate oak (Quercus robur L.) will cope with the changing climatic conditions in Europe in the near future. However, while species distribution models project expansion zones, they also identify reductions in occurrence at the dry and warm distribution margins. Whereas species distribution models primarily rely on occurrence data, tree rings––given their long-term perspective and their use in empirical models––can provide a mechanistic view of forest growth dynamics, including temporally changing climate responses. Increased climate sensitivity and growth synchrony are key dendroecological indicators of tree stress. Here, we used an unprecedented network of 150 Q. robur sites (over 3,300 trees), covering the full projected range of contracting to persistent areas across Europe, to assess the dendroecological indicators over recent decades in relation to species distribution model predictions. We reveal that oaks in areas projected to experience range contraction exhibited greater sensitivity to current growing season climatic conditions, whereas those in persistence areas responded more strongly to previous season conditions. Growth synchrony among trees was higher in the contraction areas, but showed no significant increasing trend over the last 70 years, as expected from ecotone theory. Temporal shifts in climate sensitivity were stronger for temperature and vapor pressure deficit in the persistence areas, whereas the climatic water balance gained importance in the contraction zones. These findings suggest that Q. robur growth is not yet being severely affected by climate change, and that the species is currently coping well with the climate changes, even in regions with projected range contractions, thereby challenging statistically derived scenarios of range shift based on species distribution models. Keywords: adaptation, allometry, phloemsieve element, xylem vessel Published in DiRROS: 04.11.2025; Views: 272; Downloads: 129
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5. Spatiotemporal variability of dendroecological indicators in pedunculate oak (Quercus robur L.) tree-rings across Europe in relation to species distribution modelsAndrei Popa, Jernej Jevšenak, Marcin K. Dyderski, Radosław Puchałka, Allan Buras, Ionel Popa, Martin Wilmking, Aleksandra Kalisty, Catalin Constantin Roibu, Marcin Jakubowski, 2025, original scientific article Abstract: Climate is a primary, but non-stationary, driver of tree growth. Climate change is altering the sensitivity of forest growth to water availability and temperature over time. It is considered that pedunculate oak (Quercus robur L.) will cope with the changing climatic conditions in Europe in the near future. However, while species distribution models project expansion zones, they also identify reductions in occurrence at the dry and warm distribution margins. Whereas species distribution models primarily rely on occurrence data, tree rings—given their long-term perspective and their use in empirical models—can provide a mechanistic view of forest growth dynamics, including temporally changing climate responses. Increased climate sensitivity and growth synchrony are key dendroecological indicators of tree stress. Here, we used an unprecedented network of 150 Q. robur sites (over 3300 trees), covering the full projected range of contracting to persistent areas across Europe, to assess the dendroecological indicators over recent decades in relation to species distribution model predictions. We reveal that oaks in areas projected to experience range contraction exhibited greater sensitivity to current growing season climatic conditions, whereas those in persistence areas responded more strongly to previous season conditions. Growth synchrony among trees was higher in the contraction areas, but showed no significant increasing trend over the last 70 years, as expected from ecotone theory. Temporal shifts in climate sensitivity were stronger for temperature and vapor pressure deficit in the persistence areas, whereas the climatic water balance gained importance in the contraction zones. These findings suggest that Q. robur growth is not yet being severely affected by climate change, and that the species is currently coping well with the climate changes, even in regions with projected range contractions, thereby challenging statistically derived scenarios of range shift based on species distribution models. Keywords: climate change scenarios, climate–growth relationships, climatic water balance, growth synchrony, range contraction, vapor pressure deficit Published in DiRROS: 04.11.2025; Views: 316; Downloads: 154
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6. Effects of soil and atmospheric drought on intra-annual δ13C patterns in tree ringsValentina Vitali, Jernej Jevšenak, Georg von Arx, Marina Fonti, Meisha Holloway Phillips, Rubén D. Manzanedo, Kerstin Treydte, Lorenz Walthert, Roman Zweifel, Matthias Saurer, 2025, original scientific article Abstract: None declared.Conflict of interestHigh-resolution carbon isotope ratio (δ13C) measurements of tree rings have the potential to provide seasonal environmental information. However, due to the complexity of the wood formation processes, the reliability of this method for intra-seasonal reconstruction of growing conditions remains unclear. We, therefore, investigated the intra-annual variation of δ13C in tree rings of three conifer species (Pinus sylvestris, Picea abies, Abies alba) across sites from the Swiss Alps to assess their response to seasonal variation of soil water potential (SWP) and vapour pressure deficit (VPD). Intra-annual δ13C values at a resolution of 10 points per year were assessed using laser-ablation isotope-ratio mass spectrometry. Seasonal δ13C patterns were analyzed for synchronicity across trees and species, and their correlation with on-site environmental variables was used to determine the driving factors of δ13C, to reconstruct growing season dynamics, and to estimate timings of the growth dynamics and allocation of carbon to xylem formation. The δ13C patterns showed high synchronicity between species, with characteristic maxima in wet and dry years occurring in the middle of the ring and at the end of the ring, respectively. Seasonal δ13C variations reliably reflected atmospheric dryness. Higher than normal soil dryness hindered integration of further fresh assimilates into the xylem, thus allowing the identification of species- and site-specific threshold conditions that disrupt wood formation. The δ13C of Scots pine shows the strongest correlations with VPD and SWP, making it an excellent indicator of environmental variability. Silver fir appears to integrate carbon into xylem structural material over a longer season than the other conifers, whilst Norway spruce shows more plastic site-specific responses to environmental conditions. In conclusion, we identify how atmospheric and soil drought jointly impact tree growth and intra-annual δ13C patterns across conifer species, offering valuable insights for climate reconstructions and wider applications on forest dynamics. Keywords: water availability, annual growth, laser ablation, tree-ring isotopes, carbon isotopes Published in DiRROS: 01.10.2025; Views: 364; Downloads: 189
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7. Temperature-driven shifts in spatiotemporal stability of climate-growth responses of Douglas-fir (Pseudotsuga menziesii) from the southern Baltic Sea regionMarcin Miroslav Klisz, Radosław Puchałka, Mariusz Gławenda, M. Koprowski, Roberts Matisons, Sandra Metslaid, Aleksei Potapov, Tobias Scharnweber, Eric Andreas Thurm, Rita Verbylaite, Adomas Vitas, Martin Wilmking, Jernej Jevšenak, 2025, original scientific article Abstract: The southern Baltic region spans a significant part of the European continent with its forests under significant pressure due to climate changes. The implications of these changes are crucial for both native and non-native tree species. Under future climate scenarios, most native conifer populations might lose their climatic optima in the region. In contrast, for non-native Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco), climatic conditions are expected to become optimal in the Baltic Sea region. Therefore, understanding the acclimatisation trajectory of Douglas-fir over the last century is essential to assess its potential to supplement retreating species and reduce pressure on local habitats. To study the region-wide acclimatisation in the secondary distribution, we established a network of 27 Douglas-fir tree-ring chronologies along the south Baltic Sea. We determined the spatio-temporal stability of the climate signal in tree rings and the potential coastal effect on the plasticity of the growth response. We found a region-wide trend of climate-growth relationships, with a dominant effect of the early-growth season temperatures being more pronounced for mature than young stands. Sites with higher mean annual temperatures exhibit a stronger positive temperature–growth correlation, demonstrating the sensitivity of Douglas-fir to climate warming. Douglas-fir could serve as a more heat-tolerant alternative to the declining European species of the Pinaceae family and contribute to the preservation of functionally comparable coniferous forest communities. However, forest practitioners should be aware that Douglas-fir may alter habitat conditions affecting microclimate and influencing species diversity. Keywords: climate warming, climate–growth relationships, dendroecology, Europe, tree growth Published in DiRROS: 09.07.2025; Views: 520; Downloads: 403
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8. A single-tree approach to determine climate-growth patterns of European beech and their seasonality in the species southern distribution areaRoberto Serrano-Notivoli, Jernej Jevšenak, Katarina Čufar, Nina Škrk Dolar, Angela Balzano, Jožica Gričar, Maks Merela, Klemen Novak, Peter Prislan, Martin De Luis, 2025, original scientific article Abstract: Dry and warm climate conditions in southern Europe represent clear limits for European beech (Fagus sylvatica) growth near the species southern distribution limit, but it is unclear how aridification and changes in seasonal precipitation regimes will affect these forests at the individual level. We explored climate-growth relationships and the seasonality of peak climate signals in European beech using daily climate data and a large collection of tree-ring width series from southern and southeastern Europe through Generalised Linear Mixed Models (GLMMs). In most cases we found a positive and significant influence of precipitation on tree growth, and a significant negative effect of maximum temperature. Predictions from the GLMMs revealed a positive impact of precipitation during an 88 day window from spring to early summer (mid-April to mid-July), for an average tree across our network. This critical growing time window ranged from 75 days in warmer and drier conditions, and extended up to 100 days in areas with mild temperatures and moderate summer precipitation. Maximum temperatures negatively affected trees for an average of 27 day window in summer (June-July). This period was reduced to <10 days in locations with wetter and colder summers, rising up to 45 days in sites with drier and warmer summers. The positive effect of precipitation on growth was stronger and commenced earlier in larger trees. Similarly, the negative effects of maximum temperatures were more pronounced for larger trees. The use of daily climate data and a tree-centred approach allowed for capturing critical temporal dynamics in climate-growth relationships that are often overlooked by conventional methods. These insights significantly enhance our understanding of climatic factors influencing individual beech growth at the edge of its distribution range and their seasonal variations. Keywords: European beech, daily climatic variables, precipitation, maximum temperature, tree-ring width, range limit Published in DiRROS: 06.06.2025; Views: 632; Downloads: 306
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9. Age-dependent moisture response of conifers near their cold range limitJan Tumajer, Grudd Håkan, Jernej Jevšenak, Andreas J. Kirchhefer, Francesco Marotta, Jiří Mašek, Kiara Maria Nowatzki, Nikolaus Obojes, Markus Stoffel, V. Treml, Jelena Lange, 2025, original scientific article Abstract: Growth dynamics of cold subarctic and subalpine forests are primarily sensitive to temperature, but growth responses may vary across regions or shift over tree ontogeny. Systematic assessments of spatial, interspecific, and demographic variation in temperature and moisture limitation of tree growth at cold distribution margins are essential for enhancing our understanding of how these ecosystems will evolve under climate regimes. To address this gap, we built a network of 22 tree-ring width chronologies from cold forests covering two regions (Fennoscandia, European Alps), three species (Pinus sylvestris, Pinus cembra, Larix decidua), and two age cohorts (45–100 and >150 years). We combined daily climate-growth correlations with the process-based Vaganov-Shashkin growth model to identify differences in critical growth factors between species and age cohorts. In addition, we assessed the coincidence of unusually wide and narrow tree rings with years of specific climatic anomalies. Although growing season temperature was the dominant growth-limiting factor, seasonal effects of water balance on tree growth were considerably large and varied systematically between regions, species, and particularly age cohorts. The growth of young P. sylvestris in Fennoscandia responded negatively to water balance and narrow rings coincided with wet years. In contrast, the growth of young P. cembra in the Alps was drought-limited. Old trees of all species and both age cohorts of L. decidua in the Alps showed limited sensitivity to water balance. The patterns of climate-growth responses in cohort chronologies based on tree age at the year of coring were similar to ontogenetic shifts of climate-growth responses if chronologies were based on the cambial age of individual rings. Our results stress the need to account for interspecific and demographic differences in sensitivity to climate in large-scale studies of cold forest ecosystems. Keywords: boreal forest, Larix, subalpine forest, pinus, tree ring, Vaganov-Shashkin model Published in DiRROS: 04.06.2025; Views: 633; Downloads: 313
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